@article {3856,
	title = {Excess of Deleterious Mutations around HLA Genes Reveals Evolutionary Cost of Balancing Selection},
	journal = {Molecular Biology and Evolution},
	volume = {33},
	year = {2016},
	pages = {2555{\textendash}2564},
	doi = {10.1093/molbev/msw127},
	author = {Lenz, Tobias L and Spirin, Victor and Jordan, Daniel M and Sunyaev, Shamil R}
}
